賀金生，男，北京大學城市與環境學院教授，博士生導師，生態學系副主任。973計劃首席科學家，國家杰出青年基金獲得者。主要從事化學計量生態學、草地生態系統碳循環、青藏高原高寒生態系統與全球變化等方面的研究。發表中英文論文100余篇，其中SCI論文60余篇。發表的雜志包括：New Phytologist，Oecologia，Ecology，Ecology Letters，Functional Ecology等。目前是國內外4個專業期刊的編委，其中包括重要國際刊物“Frontiers in Ecology and the Environment”和“Journal of Plant Research”。
個人簡介
學習經歷
1984.9~1988.7, 蘭州大學生物系，獲理學學士（植物學）
1988.9~1991.6,中國科學院植物研究所，獲理學碩士學位（生態學）
1996.9~1998.8, 中國科學院植物研究所，獲理學博士學位（生態學）
1999.7~2002.8, 哈佛大學進化與生物學系博士后（生物學）
2007.10~2008.1, 瑞士蘇黎世大學，The Swiss National Science Foundation Research Fellow
教育工作
1991.6~1999.11 中國科學院植物研究所，實習研究員，助理研究員，副研究員
2002.8~2008.7 北京大學生態學系，副教授
2007.1~北京大學生態學系副主任
2008.8~ 北京大學生態學系，教授
2009.3~ 西北高原生物研究所百人計劃
學術任職
《植物生態學報》編委，2004~
《生物多樣性》編委，2009~
Journal of Plant Ecology (Oxford University Press，SCI), Associate editor, 2008~
Journal of Plant Research (Springer,SCI), Editorial Board Member, 2007~2014
Frontiers in Ecology and the Environment (ESA journal,SCI), Associate editor, 2007~2014
中國植物學會植物生態學專業委員會委員，2009~
中國植物學會學術委員會委員，2009~
科學研究
研究方向
1. 植物生理生態學
2. 生態系統生態學
3. 青藏高原生態學
4. 生物多樣性的生態系統功能
當前研究領域
1、生態系統生態學：主要圍繞陸地生態系統的碳循環和氮循環, 采用實驗的手段，研究全球氣候變化影響下生態系統過程的反應。近期的研究重點是：降雨格局的改變和氮沉降如何影響溫帶草地、人工林C、N循環的主要過程。
2、植物功能屬性：通過野外植物主要生態屬性的測定，結合室內觀葉植物的解剖結構的分析，在大尺度、多生態系統類型上探討植物生態屬性的變化規律和控制因子。
3、地下生態學：通過典型生態系統地下過程的調查，探討生態系統地上和地下部分的關聯，重點探討植物地上部分得到的規律性是否可以應用到地下。
4、生物多樣性的生態系統功能：結合實驗途徑和野外調查，研究生物多樣性的生態系統功能，特別是在C, N經濟（C, N economy）中的作用。主要的實驗系統是草原生態系統。
研究項目
1、國家重點基礎研究發展規劃（973計劃）：中國陸地生態系統碳循環及其驅動機制研究（2002-2007），專題主持
2、國家自然科學基金重大研究計劃重點項目：我國溫帶草地碳庫分布、動態與氣候控制（2003-2006），專題主持
3、國家中華人民共和國國家自然科學基金委員會“創新群體”：環境生物地球化學研究（2004-2006）成員
4、國家自然科學基金重大研究計劃面上項目（90411004）：溫帶草地根系生態學的實驗研究：個體、種群和功能屬性的關聯（2005-2007），主持
5、國家自然科學基金委面上項目（30670322）：青藏高原極端環境條件下植物葉功能屬性特征及環境的修飾作用（2007-2009），主持
6、中日韓A3前瞻計劃項目（Asia 3 Foresight Program）：Quantifying and predicting terrestrial carbon sinks in East Asia: toward a network of climate change research，A3 3+2（2007-2012），參加
7、國家科技支撐計劃（2007BAC06B00）：羌塘高原高寒草甸區生態穩定性維持技術研究與示范（專題）（2007.9-2010.12），主持
8、國家中華人民共和國國家自然科學基金委員會面上項目（30870381）：我國主要溫帶草地優勢植物葉片、根系和土壤C：N：P化學計量學的關聯（2009-2011），主持
9、歐盟項目（BACCARA）：Biodiversity And Climate Change，A Risk Analysis（2009-2011），專題3
10、科技部國際合作（Sino-EU 0916）：Risk analysis of biodiversity change：a case study in a subtropical forest，China（2009-2012），主持
11、基金委創新群體項目（31021001）：中國陸地植被的時空格局與生態功能（2011-2013），核心成員
12、"全球變化研究國家重大科學研究計劃"項目（2010CB950600）：中國陸地生態系統碳源匯特征及其全球意義（2010-2013），研究骨干
13、國家杰出青年科學基金（31025005）：植物功能屬性與化學計量生態學（2011-2014），主持
14、全球變化研究國家重大科學研究計劃項目（2014CB954004）（973）“土壤系統碳動態、機制及其對全球變化的響應”（2014-2018），首席科學家
主要榮譽
1998年，中科院地奧獎學金"一等獎, 博士論文"神農架林區地區米心水青岡林和銳齒槲櫟林的格局和過程"
2003年，教育部自然科學一等獎：中國陸地生態系統碳循環與植被生產力的研究，第3完成人
2004年，國家自然科學二等獎：中國陸地生態系統生產力和碳循環的研究，第5完成人
2006年，北京大學“保潔”獎
2007年，Sino-Swiss Science and Technology Cooperation Research Fellowship, Swiss Federal Institute of Technology (ETH Zurich), Swiss National Science Foundation
2007年，北京大學優秀班主任三等獎
2010年，"國家杰出青年科學基金" 獲得者
2014年，北京大學“保潔”獎
主講課程
代表論著
Jing X, Sanders NJ, Shi Y, Chu HY, Classen AT, Zhao K, Chen LT, Jiang YX, He J-S* 2015.The links between ecosystem multifunctionality and above- and belowground 生物多樣性 are mediated by climate.Nature Communications6: 8159.
Song WM, Wang H, Wang GS, Chen LT, Jin ZN, Zhuang QL, He J-S*. 2015.甲烷 emissions from an alpine wetland on the Tibet Plateau: Neglected but vital contribution of non-growing season.Journal of Geophysical Research-Biogeosciences: doi:10.1002/2015JG003043.
Jin ZN, Zhuang QL, He J-S, Zhu XD, Song WM 2015.Net exchanges of 甲烷 and carbon dioxide on the Qinghai-Tibet Plateau from 1979 to 2100.Environmental Research Letters: doi:10.1088/1748-9326/10/8/085007.
Niu KC, He J-S, Zhang ST, Lechowicz MJ 2015.Grazing increases functional richness but not functional divergence in Tibet alpine 草甸 plant communities.Biodiversity and Conservation: doi: 10.1007/s10531-015-0960-2.
Chen LT, Flynn DFB, Zhang XW, Gao XL, Lin L, Luo J, Zhao CM* 2015.Divergent patterns of foliar δ13C and δ15N in Quercus aquifolioides with an altitudinal transect on the Tibet Plateau: an integrated study based on multiple key leaf functional traits.Journal of Plant Ecology8: 303-312.
Chen LT, Flynn DFB, Jing X, Kühn P, Scholten T, He J-S 2015.A comparison of two methods for quantifying soil organic carbon of alpine grasslands on the Tibet 高原PLoS ONE10: e0126372.
Mi ZR, Huang YY, Gan HJ,Zhou WJ, Flynn DFB,He J-S* 2015.Leaf P increase outpaces leaf N in an Inner Mongolia grassland over 27 years.Biology Letters 11: 20140981.
Ding S, Xu Y, Wang Y, He Y,Hou J, Chen L,He J-S 2015.Distributions of 丙三醇 dialkyl glycerol tetraethers in surface soils ofQinghai–Tibet Plateau: implications of GDGT-based proxies in cold and dryregions.Biogeosciences Discussion 12: 481-513.
Zhang ZJ,Yan YJ,Tian Y, LiJS,He J-S, Tang ZY* 2015.Distribution and conservation of orchid 物種 richness in China.Biological Conservation 181: 64-72.
Eichenberg D, Trogisch S,Huang YY,He J-S, Bruelheide H 2014.Shifts in community leaf functional traits are related to litter decompositionalong a secondary forest succession series in subtropical China.Journal of Plant Ecology DOI:10.1093/jpe/rtu021: .
Wang YH, Liu HY, Chung H,Yu LF, Mi ZR, Gen.G Y, Jing X, Wang SP, Zeng H, Cao GM, Zhao XQ,He J-S* 2014. Non-growing-season soilrespiration is controlled by freezing and thawing processes in the summermonsoon-dominated Tibet alpine grassland.Global Biogeochemical Cycles28: 1081-1095.
Yang XX, Yang Y, Ji CJ,Feng T, Shi Y, Lin L, Ma JJ,He J-S*2014. Large-scale patterns of stomatal traits in Tibet and Mongoliangrassland 物種Basic and Applied Ecology 15: 122-132.
Shi Y, Wang YH, Ma YL, MaWH, Liang CZ, Flynn DFB, Schmid B, Fang JY,He J-S* 2014. Field-based observations of regional-scale, temporalvariation in net primary production in Tibet alpine grasslands.Biogeosciences11: 2003-2016.
Baumann F*, Schmidt K,D?rfer C,He J-S, Scholten T, Kühn P2014. Pedogenesis, permafrost, substrate and topography: Plot and landscapescale interrelations of 風化作用 processes on the central-eastern Tibet高原Geoderma 226-227: 300-316.
Geng Y, Wang L, Jin DM, LiuHY,He J-S* 2014. Alpine climatealters the relationships between leaf and root morphological traits but notchemical traits.Oecologia 175: 445-455.
Jing X, Wang YH, Chung H,Mi ZR, Wang SP, Zeng H,He J-S*2014. No 溫度 acclimation of soil extracellular enzymes to experimentalwarming in an alpine grassland ecosystem on the Tibet Plateau.Biogeochemistry117: 39-54.
Chen LT, Niu KC, Wu Y, GengY, Mi ZR, Flynn DFB,He J-S* 2013.UV radiation is the primary factor driving the variation in leaf phenolicsacross Chinese grasslands.Ecology and Evolution 3: 4696-4710.
Jin DM, Ma JJ, Ma WH, LiangCZ, Shi Y andHe J-S* 2013. Legumesin Chinese natural grasslands: 物種, biomass, and distribution.Rangeland Ecology & Management 66: 648-656.
Du EZ, Zhou Z, Li P, Hu XY,Ma YC, Wang W, Zheng CY, Zhu JX,He J-S,Fang JY* 2013. NEECF: a project of nutrient enrichment experiments in China’sforests.Journal of Plant Ecology 6: 428-435.
HeJ-S*,Muraoka H, Son Y, Fang JY 2013. Carbon patterns and processes in East Asianecosystems: multi-scale approaches (Editorial).Journal of Plant Ecology6: 323-324.
Ma YL, Geng Y, Huang YY,Shi Y, Niklaus P, Schmid B,He J-S*2013. Effect of clear-cutting silviculture on soil respiration in a subtropicalforest of China.Journal of Plant Ecology 6: 335-348.
Yu LF, Wang H, Wang GS,Song WM, Huang Y, Li S-G, Liang NS, Tang YH,He J-S* 2013. A comparison of 甲烷 emission measurements usingeddy covariance and manual and automated chamber-based techniques in TibetPlateau alpine wetland.Environmental Pollution 181: 81-90.
Liu C, Wang XP, Wu X, DaiS,He J-S, Yin WL* 2013. Relativeeffects of phylogeny, biological characters and environments on leaf traits inshrub biomes across central Inner Mongolia, China.Journal of Plant Ecology 6: 220-231.
Jin ZN*, Zhuang QL,He J-S, Luo TX, Shi Y 2013. Phenologyshift from 1989 to 2008 on the Tibet Plateau: an analysis with aprocess-based soil physical model and remote sensing 數據Climatic Change 119: 435-449.
Shi Y, Ma YL, Ma WH, LiangCZ, Zhao XQ, Fang JY,He J-S* 2013.Large scale patterns of forage yield and quality across Chinese grasslands.Chinese Science Bulletin 58: 1187-1199.
Dorfer C*,Kuhn P,Baumann F,He J-S,Scholten T 2013. Soil organiccarbon pools and stocks in permafrost-affected soils on the Tibet Plateau.PLoS ONE 8: e57024.
Huang N*,He J-S, Niu Z 2013. Estimating thespatial pattern of soil respiration in Tibet alpine grasslands using LandsatTM images and MODIS 數據Ecological Indicators 26: 117-125.
HeJ-S 2012.Carbon cycling of Chinese forests: From carbon storage, 動力學 to models.Science China Life Science 55: 188-190.
Geng Y, Wang ZH, Liang CZ,Fang JY, Frank B, Kühn P, Scholten T,HeJ-S* 2012. Effect of geographical range size on plant functional traits andthe relationships between plant, soil and climate in Chinese grasslands.Global Ecology and 生物地理學 21: 416–427.
Shi Y, Baumann F, Ma YL,Song C, Kuehn P, Scholten T,He J-S*.2012. Organic and inorganic carbon in the topsoil of the Mongolian and Tibetgrasslands: pattern, control and implications.Biogeosciences 9:2287–2299.
Geng Y,Wang YH,Yang K,WangSP,Zeng H,Baumann F, Kuehn P, Scholten T,HeJ-S* 2012. Soil Respiration in Tibet Alpine Grasslands: BelowgroundBiomass and Soil Moisture, but Not Soil 溫度, Best Explain theLarge-Scale Patterns.PLoS ONE 7: e34968.
Ma JJ, Ji CJ, Han M, ZhangTF, Yan XD, Hu D, Zeng H,He J-S*2012. Comparative analyses of leaf anatomical characteristics of dicotyledonous物種 in Tibet and Inner Mongolian grasslands.Science China Life Sciences 42: 158-172.
Tao S*, Wang WT, Liu WX,左姓 Q, Wang XL, Wang R, Wang B, Shen GF, Yang YH,He J-S 2011. Polycyclic aromatic hydrocarbons and organochlorinepesticides in surface soils from the Qinghai-Tibet plateau.Journal of Environmental Monitoring 13: 175–181.
Chu ZY, Lu YJ, Chang J*,Wang M, Jiang H,He J-S, Peng CH*,Ge Y 2011. Leaf respiration/photosynthesis relationship and variation: aninvestigation of 39 woody and herbaceous species in east subtropical China.Trees-Structureand Function 25: 301-310.
Ma WH,He J-S*, Yang YH, Wang XP, Liang CZ, Anwar M, Zeng H, Fang JY,Schmid B 2010. Environmental factors co-vary with plant diversity-productivityrelationships among Chinese grassland sites.Global Ecology and 生物地理學 19: 233–243.
HeJ-S*, WangX, Schmid B, Flynn DFB, Li XF, Reich PB, Fang JY 2010. Taxonomic identity,phylogeny, climate and soil fertility as drivers of leaf traits across Chinesegrassland biomes.Journal of Plant Research 123: 551-561.
Ma WH, Liu ZL, Wang ZH,Wang W, Liang CZ, Tang YH,He J-S,Fang JY* 2010. Climate change alters interannual variation of grasslandaboveground productivity: evidence from a 22-year measurement series in theInner Mongolian grassland.Journal of Plant Research 123:509-517.
Baumann F,He J-S,Schmidt K, Kühn P, Scholten T. 2009. Pedogenesis, permafrost, and soil moistureas controlling factors for soil 氮 and carbon contents across the Tibet高原Global Change Biology 15: 3001–3017
Yang YH, Fang JY, Smith P,Tang YH, Chen AP, Ji CJ, Hu HF, Rao S, Tan K,He J-S. 2009. Changes intopsoil carbon stock in the Tibet grasslands between the 1980s and 2004.Global Change Biology 15: 2723–2729
He J-S, Wang XP, Flynn DF, WangL, Schmid B, Fang JY. 2009. Taxonomic, phylogenetic and environmental tradeoffsbetween leaf 生產率 and persistence.Ecology 90: 2779-2791
He J-S, Wang L, Flynn DFB, WangX, Ma W, Fang J. 2008. Leaf 氮: 磷 stoichiometry across Chinesegrassland biomes.Oecologia155, 301-310
He J-S, Fang JY, Wang ZH, Guo DL,Flynn DFB, Geng Z. 2006. Stoichiometry and large-scale patterns of leaf carbonand 氮 in the grassland biomes of China.Oecologia,149:115-122
He J-S, Wang ZH, Wang XP, SchmidB, Zuo W, Zhou M, Zheng CY, Wang MF, Fang JY, 2006. A test of the generality ofleaf trait relationships on the Tibet 高原New Phytologist, 170: 835-848
Balvanera P, Pfisterer AB,Buchmann N,He J-S, Nakashizuka T, Raffaelli D, Schmid B. 2006.Quantifying the evidence for biodiversity effects on ecosystem functioning andservices.Ecology Letters, 99:1146–1156
He J-S, Flynn DFB, Wolfe-BellinK, Fang J, Bazzaz FA. 2005. CO and 氮, but not populationdensity, alter the size and C/N ratio ofPhytolacca americana seeds.Functional Ecology, 19: 437-444
He J-S, Wolfe-Bellin KS, SchmidB, Bazzaz FA. 2005. 密度 may alter diversity-生產率 relationships inexperimental plant communities.Basicand Applied Ecology, 6: 505-517
He J-S, Wolfe-Bellin KS, BazzazFA. 2005. Leaf-level physiology, biomass, and reproduction ofPhytolaccaamericana under conditions of elevated CO and alteredtemperature regimes.International Journal of Plant Sciences, 166: 615-622
He J-S, Bazzaz FA. 2003.密度dependent responses of reproductive allocation to elevated atmosphericCO inPhytolacca americana.New Phytologist, 157: 229-239
He J-S, Bazzaz FA, Schmid B.2002. Interactive effects of diversity, nutrients and elevated COon experimental plant communities.Oikos,97: 337-348
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